The recent events that unfolded in the UK with the nerve agent poisoning of the former Russian spy Sergei Skripal and his daughter on a bench in Salisbury is putting the secret labs of the Russians that research and manufacture these weapons in the spotlight.

During his last run for the presidency, in 2012, Russian leader Vladimir Putin startled U.S. military experts with a mysterious pledge to develop novel kinds of weapons to counter the West’s technological edge. Armies of the future, he said, would need weapons “based on new physical principles” including “genetic” and “psychophysical” science.

“Such high-tech weapons systems will be comparable in effect to nuclear weapons,” Putin said in an essay published in Rossiyskaya Gazeta, the Russian government’s newspaper of record, “but will be more ‘acceptable’ in terms of ­political and military ideology.”

Since the start of Putin’s second term, a construction boom has been underway at more than two dozen institutes that were once part of the Soviet Union’s biological and chemical weapons establishment, according to Russian documents and photos compiled by independent researchers. That expansion, which includes multiple new or refurbished testing facilities, is particularly apparent at secret Defense Ministry laboratories that have long drawn the suspicions of U.S. officials over possible arms-treaty violations.

“The big question is, why are they doing this?” said Raymond Zilinskas, a chemical and biological weapons expert with the James Martin Center for Nonproliferation Studies in Monterey, Calif. In a newly released book, “Biosecurity in Putin’s Russia,” ­Zilinskas and co-author Philippe Mauger analyze hundreds of contract documents and other records that show a surge in Russian research interest in subjects ranging from genetically modified pathogens to nonlethal chemical weapons used for crowd control.

During the Cold War, Soviet leaders spent vast sums to create weaponized versions of 11 different pathogens — including the microbes that cause anthrax, smallpox and the plague — while also experimenting with genetically altered strains. They created new classes of chemical toxins, such as Novichok, reportedly used in the attempted assassination of former Russian spy Sergei Skripal and his daughter, Yulia, in Salisbury, England.

A fourth-generation nerve agent more deadly than VX, Novichok is the stuff of legend. Russia denies that it ever researched or manufactured such nerve agents, but it arrested a former Soviet weapons scientist on charges of divulging state secrets after he published details about Soviet Novichok production in newspaper articles and a memoir.

Even after the fall of the former Soviet Union, the Russians didn’t allow inspectors inside their former chemical warfare research labs. Some thought the shape of the labs was part of the reason but many in the intelligence field believed that they were hiding something.

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